Does anyone know what these figures are doing with their feet? I keep seeing it described as "wrestling," but I've never found any explanation, and I've found versions of this image everywhere.

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(BL, MS Add 42130, f. 152v; Paris, Bibl. Sainte-Geneviève, MS 0143, f. 001; Frauenfeld, Kantonsbibl. Thurgau Y 24, f. 12; Bod Lib, MS Douce 6, f. 117r)
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